Structure of the LDL receptor extracellular domain at endosomal pH

Rudenko, G; Henry, L; Henderson, K; Ichtchenko, K; et al

HERO ID

4683798

Reference Type

Journal Article

Year

2002

HERO ID 4683798
In Press No
Year 2002
Title Structure of the LDL receptor extracellular domain at endosomal pH
Authors Rudenko, G; Henry, L; Henderson, K; Ichtchenko, K; et al
Journal Science
Volume 298
Issue 5602 (Dec 20
Page Numbers 2353-2358
Abstract   The low-density lipoprotein receptor mediates cholesterol homeostasis through endocytosis of lipoproteins. It discharges its ligand in the endosome at pH < 6. In the crystal structure at pH = 5.3, the ligand-binding domain (modules R2 to R7) folds back as an arc over the epidermal growth factor precursor homology domain (the modules A, B, beta propeller, and C). The modules R4 and R5, which are critical for lipoprotein binding, associate with the beta propeller via their calcium-binding loop. We propose a mechanism for lipoprotein release in the endosome whereby the beta propeller functions as an alternate substrate for the ligand-binding domain, binding in a calcium-dependent way and promoting lipoprotein release.   The low-density lipoprotein receptor mediates cholesterol homeostasis through endocytosis of lipoproteins. It discharges its ligand in the endosome at pH < 6. In the crystal structure at pH = 5.3, the ligand-binding domain (modules R2 to R7) folds back as an arc over the epidermal growth factor precursor homology domain (the modules A, B, beta propeller, and C). The modules R4 and R5, which are critical for lipoprotein binding, associate with the beta propeller via their calcium-binding loop. We propose a mechanism for lipoprotein release in the endosome whereby the beta propeller functions as an alternate substrate for the ligand-binding domain, binding in a calcium-dependent way and promoting lipoprotein release.
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Dupe Override No
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Keyword Lipids; Proteins; Cholesterol; 2002)